Key E: Stratified Foliose Cyanolichens

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1a Thallus with veins on the lower surface (occasionally the veins ± obscure); lower cortex absent (appearing dull and fibrous under lens)

2a Medulla white; lower surface never orange

Peltigera

2b Medulla orange; lower surface orange with brown veins with brown veins

Solorina crocea

1b Thallus without veins on the lower surface; lower cortex present or not

3a Thallus minutely foliose or subfoliose; lobes < 2 mm wide

4a On bark or wood

5a Cortex or medulla P+O; thallus with definite lobes

Pannaria

5b Cortex and medulla P-; thallus with definite lobes or nearly squamulose

Fuscopannaria

(Parmeliella is also keyed under Fuscopannaria. Koerberia biformis, a minute blackish-olive species on trees in Cal, would also key here.)

4b On rock, mosses, or humus over rock

6a On mosses or humus over rock or bark, sometimes directly on rock; thallus brown or bluish brown; lobes never longitudinally ridged

7a Thallus brown or dark brown; lobes isidiate to lobulate; spores 2-4 celled

Massalongia

7b Thallus gray or brownish gray; lobes isidiate becoming sorediose; spores simple

Fuscopannaria (including Moelleropsis)

6b On rock; thallus blackish, gray green, or olive colored; lobes often longitudinally ridged or striate

8a Lower surface dark

Placynthium (not treated)

8b Lower surface pale

9a Isidia and lobules lacking; apothecia usually present. Arctic s to s WA, rare in PNW

Vestergrenopsis elucina

9b Isidia and/or lobules present; apothecia occasionally present

10a Isidia very slender, cylindrical to + flattened; lobes very thin, elongate, fan-tipped, and tightly appressed; apothecia margins thalline; rare w Cas, more common northward

Vestergrenopsis isidiata

10b Isidia present or not, but with abundant narrow lobules, the lobules mostly flattened (some cylindrical) and arising from fine divisions of the lobe margins, often becoming densely shingled on the upper surface; lobes thicker, shorter, and less tightly appressed than the preceding species, generally fan-tipped in part; apothecia margins variable, thalline or with a false proper margin

Koerberia sonomensis

3b Thallus medium to large foliose; lobes > 2 mm wide

11a Lower surface with discrete white or yellow spots or pores (cyphellae or pseudocyphellae) < 2 mm diameter

12a Lower surface with broad (generally 0.5-2 mm) whitish craters (cyphellae)

Sticta

12b Lower surface with small (generally < 0.5 mm) white or yellow spots (pseudocyphellae or papillae)

13a Spots on lower surface white or yellow, concave to slightly convex (pseudocyphellae); apothecia, if present, on the upper surface; upper surface dark brown, brown, gray, or greenish gray

Pseudocyphellaria

13b Spots on lower surface white, raised (papillae); apothecia on the lower surface of the lobe tips; upper surface brown togray brown

Nephroma resupinatum

11b Lower surface lacking discrete white or yellow spots or pores, but in some spp with whitish patches > 2 mm diameter

14a Lobes large, usually > 1 cm broad; lower surface with fine tomentum interrupted by naked, white patches; upper surface weakly undulating or with a network of prominent ridges

Lobaria

14b Lobes smaller, usually < 1 cm broad; lower surface without naked, white patches or without tomentum; upper surface weakly or not at all ridged

15a Apothecia on lower surface of lobe tips; thallus brown to gray brown; soredia present or not

Nephroma

15b Apothecia absent or not on lower surface; thallus gray, brownish gray, olive, greenish gray or yellowish; soredia or lobules present; rare spp

16a Lobules or apothecia present

17a Lobules present, containing green algae, thallus otherwise with a blue-green photobiont; soredia absent

Lobaria oregana (blue-green photomorph)

17b Lobules present or not; apothecia present, the whole thallus containing cyanobacteria

18a Lower surface with a felty mat of blue-black rhizines and tomentum; thallus P-. Arctic-alpine, Alas and OP; rare

Coccocarpia erythroxyli

18b Lower surface with whitish or bluish rhizohyphae; thallus P+O (eriodermin)

[Erioderma pedicellatum]

16b Lobules absent; soredia present and containing blue-green photobiont

19a Soredia laminal; lower surface bare; upper surface yellowish tinged when dry, bluish gray when moist

Nephroma occultum

19b Soredia marginal; lower surface tomentose; upper surface grayish, not tinged with yellow

20a Upper surface with stiff erect hairs (use lens); thallus P+O

Erioderma sorediatum

20b Upper surface with matted cobwebby hairs, never with stiff erect hairs; thallus P-

Leioderma sorediatum